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The petiolate leaves are dentate and often sticky. The lower leaves are opposite, ovate, or often heart-shaped. They are distinguished technically by the fact that the ray florets are sterile, and by the presence on the disk flowers of a pappus that is of two awn-like scales that are caducous . Some species also have additional shorter scales in the pappus, and one species lacks a pappus entirely. Another technical feature that distinguishes the genus more reliably, but requires a microscope to see, is the presence of a prominent, multicellular appendage at the apex of the style. Further, the florets of a sunflower are arranged in a natural spiral.